It surely will...as long as the water flows it's plenty enough to be on the safe side.

It sort of depends on what you mean by enough. The AMS rads, the Kryos and I guess Aquacomputer Titan GPU blocks? are all relatively restrictive components compared to other brands.
I've never seen accurate figures for the Kryos CPU block but relative flow to other blocks puts it around the swiftech CPU blocks roughly.
The AMS rads are around 0.8 PSI drop
An AC Titan block is around 2.5 PSI drop

A D5 with an after market top produces enough pressure (4.7ish PSI) to just push 1 GPM through 2 AC Titan blocks in series. Any more than that and the flow rate will be lower. How much lower is much harder to work out, but it could be significantly less.
That is not a world ending big deal because in general lower flow rates only mean small component temp (vs water) increases on each block. It can make bleeding air out of the system more difficult and also means that when blocks are connected in series that the water warms more after flowing through each block, which can then mean each block following another gets progressively warmer. Low water flow also decreases radiator efficiency.

All that can almost certainly be avoided by running the GPU blocks in parallel. That will reduce the overall loop restriction to a level that the pump should have no problem pushing 1 GPM through.

Citation de "Scott-S6"

...or with using lower resistance blocks....

However, the waterblocks with lower resistance from other manufacturers are not as effective and good looking as the Titan/780 blocks from Aqua Computer.
I myself operate two GTX 570 (SLI) with waterblocks in a parallel loop setting for nearly two years now and have no problems at all.
I think it is exaggerated, prejudice and almost superstition to believe that a parallel flow arrangement is not as save as an assembly in series.